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الخميس، 14 أبريل 2016

1 Introduction

Migration is commonly thought of as a mechanism by which organisms

avoid unfavorable environments for more auspicious ones. Migration can

be the seasonal movement of organisms from place to place or a more

permanent dispersal from one region to another; either way, changing

environmental conditions provide the impetus. But why do some

organisms migrate and not others? How do the benefits of migration

compare with the metabolic costs and hazards of the journey? Do

differences between terrestrial and marine species in their interface with

the environment have different effects on migration? What are the trade-
offs between migration and evolutionary adaptation to local conditions?

Different species respond to environmental changes in different ways,

partly based on their individual adaptations and range of tolerance. This

phenomenon is most easily seen when comparing terrestrial and marine

organisms, which have different relations to their environment. Terrestrial

organisms are surrounded by low-density atmosphere and expend

considerable energy moving and keeping themselves upright; marine

organisms, in contrast, are surrounded by seawater, which has a density

close to their own, making support and movement much less costly

(Sverdrup et al. 2005). Thus, migration is likely to be more metabolically

costly to a terrestrial animal than a marine one, all other things being

equal. The factors affecting migration of land and sea species may

therefore differ considerably, and one geologic episode may affect the two

realms differently.

Terrestrial and marine organisms also differ in the environmental factors

that affect them the most. Temperature and rainfall are major determinants

on the land, but salinity is one of the most important in the sea. The

difference in the salt content of seawater and bodily fluids affects the

transport of water molecules through cell membranes in a process called

osmosis; changes in salinity can have disastrous effects on the

physiological balance of an organism. Temperature variation affects

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